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From 2006 to 2009, he served as an Alaskan governor.

In 2016, Paulson portrayed real life prosecutor Marcia Clark in the first season of the anthology series American Crime Story, subtitled The People v. She lived in Queens and Gramercy Park, before settling in Park Slope, Brooklyn.She spent her summers in Florida visiting her father.She also starred in a number of independent films, and had a leading role on the ABC comedy series Cupid in 2009.She later starred in the independent drama Martha Marcy May Marlene (2011), and received Primetime Emmy Award and Golden Globe Award nominations for her performance as Nicolle Wallace in the HBO film Game Change (2012).She starred as the main character in the short-lived NBC series Leap of Faith.

In 2004, she had a supporting role in the ABC series The D. In theater productions, Paulson starred in a revival of The Glass Menagerie on Broadway and appeared Off-Broadway in Killer Joe, and Colder Than Here.Paulson starred as Mary Epps in the 2013 historical drama film 12 Years a Slave and as Abby Gerhard in the 2015 drama film Carol.Both films received widespread critical acclaim and were nominated for multiple Academy Awards.Paulson's film credits include Down with Love, What Women Want, The Other Sister, Levitation, and Serenity.In the 2006–07 television season, Paulson co-starred in NBC's Studio 60 on the Sunset Strip, playing Harriet Hayes, one of the stars of the show-within-a-show.She returned for the fifth season, subtitled Hotel, portraying the role of Hypodermic Sally.